When Morgan hybridised yellow-bodied, white-eyed females to brown-bodied, red-eyed males and intercrossed their F₁ progeny, the F₂ ratio deviated very significantly from the 9:3:3:1 ratio. This can be attributed to the fact that:

Morgan coined the term ______ to describe this physical association of genes on a chromosome and the term _________ to describe the generation of non-parental gene combinations.

The number of linkage groups in a plant with a normal diploid number of chromosomes in a somatic cell as 16 will be:
1. 16
2. 8
3. 4
4. It is impossible to tell from the information provided.